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Deadly giant bugs have begun to rampage through Edo Japan, and the Mushibugyou Protection Office must exterminate the threat. With the body count rising, the group requests aid from famed samurai Genjuurou Tsukishima, but due to an injury the man sends his inept-but-determined son Jinbei in his stead. Almost killed in his first encounter with the creatures, Jinbei is saved by his new colleagues of the Mushibugyou: fiery ninja Hibachi, violent warrior Koikawa, young onmyouji Ichinotani, and the enigmatic Mugai. Now, the group must learn to work together to overcome the many-legged menace!